sevinПОВОЛЖСКИЙ ЭКОЛОГИЧЕСКИЙ ЖУРНАЛPovolzhskiy Journal of Ecology1684-73182541-8963Федеральное государственное бюджетное учреждение науки Институт проблем экологии и эволюции им. А. Н. Северцова РАН; Товарищество научных изданий КМК.10.35885/1684-7318-2026-2-131-154sevin-751Research ArticleСтатьиПроблемы таксономии, распространение и природоохранный статус мышовок Sicista (Rodentia, Dipodoidea) ПоволжьяProblems of taxonomy, distribution and conservation status of Sicista (Rodentia, Dipodoidea) birch mice of the Volga regionhttps://orcid.org/0000-0003-0632-4949БаскевичМ. И.BaskevichM. I.

Баскевич Марина Исаковна

Лаборатория микроэволюции млекопитающих

119071 , г. Москва , Ленинский проспект , д. 33

Marina I. Baskevich

Laboratory of Mammalian Microevolution

33 Leninsky Prosp. , Moscow 119071

mbaskevich@mail.ruhttps://orcid.org/0000-0001-7698-5887ХляпЛ. А.KhlyapL. A.

Хляп Людмила Айзиковна

119071 , г. Москва , Ленинский проспект , д. 33

Liudmila A. Khlyap

33 Leninsky Prosp. , Moscow 119071

khlyap@mail.ruhttps://orcid.org/0000-0002-9629-7579ОпаринМ. Л.OparinM. L.

Опарин Михаил Львович

119071 , г. Москва , Ленинский проспект , д. 33

Mikhail L. Oparin

33 Leninsky Prosp. , Moscow 119071

oparinml@mail.ruhttps://orcid.org/0000-0002-2106-3989БогдановА. С.BogdanovA. S.

Богданов Алексей Станиславович

119334 , г. Москва , ул. Вавилова , д. 26

Alexey S. Bogdanov

26 Vavilova St ., Moscow 119334

bogdalst@yahoo.comИнститут проблем экологии и эволюции им. А. Н. Северцова РАНРоссияA. N. Severtsov Institute of Ecology and Evolution , Russian Academy of SciencesRussian FederationИнститут биологии развития им. Н. К. Кольцова РАНРоссияKoltzov Institute of Developmental Biology , Russian Academy of SciencesRussian Federation20260908202602131154Copyright © Баскевич М.И., Хляп Л.А., Опарин М.Л., Богданов А.С., 20262026Баскевич М.И., Хляп Л.А., Опарин М.Л., Богданов А.С.Baskevich M.I., Khlyap L.A., Oparin M.L., Bogdanov A.S.Данная работа распространяется под лицензией Creative Commons Attribution 4.0.This work is licensed under a Creative Commons Attribution 4.0 License.https://sevin.elpub.ru/jour/article/view/751

Впервые на основе анализа собственных и литературных хромосомных и молекулярных данных, изучения музейных коллекций и региональных Красных книг обобщены сведения о таксономическом разнообразии, особенностях распространения и категориях статуса мышовок, рода Sicista на территории Поволжья. Подтверждена важная роль хромосомных и молекулярных маркеров в уточнении таксономического статуса видов и форм Sicist a и их размещения в различных регионах Поволжья. Выявлен градиент в разнообразии рода, представленном видами-двойниками и формами неясного таксономического ранга Sicista групп betulina и subtilis, в целом совпадающий со сменой в направлении с севера на юг природных зон в Поволжье. Из Красных книг регионов Поволжья и Приложения к ним извлечены данные о категории статуса охраняемых видов и форм мышовок и отчасти показана роль ООПТ Поволжья в сохранении редких видов и форм Sicista .

For the first time, information on the taxonomic diversity, distribution features and categories of status of birch mice of the genus Sicista in the Volga region was summarized based on an analysis of our own and literary chromosomal and molecular data, a study of museum collections and regional Red Data Books. The important role of chromosomal and molecular markers in clarifying the taxonomic status of Sicista species and forms and their location in various districts of the Volga region has been confirmed. A gradient was identified in the biodiversity of the genus, represented by sibling species and forms of an unclear taxonomic rank Sicista of the betulina and subtilis groups, generally coinciding with the north-to-south change of the natural zones in the Volga region. Data on the status category of protected species and forms of mice were extracted from the Red Data Books of the Volga regions and the Appendices to them, and the role of protected areas in the Volga region in the preservation of rare species and forms of Sicista was partly shown.

Sicistaхромосомные маркерымолекулярные ( cytbIRBP ) маркерывиды-двойникипредполагаемые видыКрасные книгикатегории статусаПоволжьеSicistachromosomal markersmolecular ( cytbIRBP ) markerssibling speciesproposed speciesRed Data booksstatus categoriesVolga regionРабота выполнена в рамках государственных заданий Института проблем экологии и эволюции им. А. Н. Северцова РАН (проекты № FFER 2021-0003, 2024/2028 (тема № 1022040700412-9-1.6.12)) и Института биологии развития им. Н. К. Кольцова РАН (проект № 0088-2024-0011).This work was performed within the framework of the state assignment of the A. N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences (Projects no. FFER-2021-0003 and 2024/2028 (No. 1022040700412-9-1.6.12)) and Koltzov Institute of Developmental Biology, Russian Academy of Sciences (project No. 0088-2024-0011).ReferencesAbrakhina I. B., Osipova V. B., Tsarev G. N. Pozvonochnye zhivotnye Ul'yanovskoi oblasti [Vertebrates of the Ulyanovsk Region]. Ulyanovsk, Simbirskaya kniga, 1993. 246 p. (in Russian).Abrakhina I. B., Osipova V. B., Tsarev G. N. Pozvonochnye zhivotnye Ul'yanovskoi oblasti [Vertebrates of the Ulyanovsk Region]. Ulyanovsk, Simbirskaya kniga, 1993. 246 p. (in Russian).Andreychev A. V., Kuznetsov V. A. About change of the mammals list and categories in the Red book of Republic Mordovia. Proceedings of the Samara Scientific Center of the Russian Academy of Sciences, 2012, vol. 14, no. 5, pp. 163–167 (in Russian).Andreychev A. V., Kuznetsov V. A. About change of the mammals list and categories in the Red book of Republic Mordovia. Proceedings of the Samara Scientific Center of the Russian Academy of Sciences, 2012, vol. 14, no. 5, pp. 163–167 (in Russian).Appendix 2. List of species requiring special monitoring of their status in the wild within the Nizhny Novgorod region. In: Red Book of the Nizhni Novgorod Region. Vol. 1. Animals. 3rd ed., rev. and expanded. Nizhni Novgorod, Ecotsenter “Dront”, 2026, pp. 402 (in Russian).Appendix 2. List of species requiring special monitoring of their status in the wild within the Nizhny Novgorod region. In: Red Book of the Nizhni Novgorod Region. Vol. 1. Animals. 3rd ed., rev. and expanded. Nizhni Novgorod, Ecotsenter “Dront”, 2026, pp. 402 (in Russian).Baskevich M. I. Northern birch mouse Sicista betulina Pallas, 1779. In: Lissovsky A. A., Stakheev V. V., Saveljev A. P., Ermakov O. A., Smirnov D. G., Glazov D. M., Obolenskaya E. V., Sheftel B. I., Titov S. V., eds. Atlas of Mammal Distribution in the European Part of Russia. Moscow, KMK Scientific Press, 2025a, pp. 298 (in Russian).Baskevich M. I. Northern birch mouse Sicista betulina Pallas, 1779. In: Lissovsky A. A., Stakheev V. V., Saveljev A. P., Ermakov O. A., Smirnov D. G., Glazov D. M., Obolenskaya E. V., Sheftel B. I., Titov S. V., eds. Atlas of Mammal Distribution in the European Part of Russia. Moscow, KMK Scientific Press, 2025a, pp. 298 (in Russian).Baskevich M. I. Strans’s birch mouse Sicista strandi Formosov, 1931. In: Lissovsky A. A., Stakheev V. V., Saveljev A. P., Ermakov O. A., Smirnov D. G., Glazov D. M., Obolenskaya E. V., Sheftel B. I., Titov S. V., eds. Atlas of Mammal Distribution in the European Part of Russia. Moscow, KMK Scientific Press, 2025b, pp. 299 (in Russian).Baskevich M. I. Strans’s birch mouse Sicista strandi Formosov, 1931. In: Lissovsky A. A., Stakheev V. V., Saveljev A. P., Ermakov O. A., Smirnov D. G., Glazov D. M., Obolenskaya E. V., Sheftel B. I., Titov S. V., eds. Atlas of Mammal Distribution in the European Part of Russia. Moscow, KMK Scientific Press, 2025b, pp. 299 (in Russian).Baskevich M. I., Oparin M. L. On new finding of the birch mouse Sicista strandi (Rodentia, Dipodoidea) specifying the northeastern boundary of the species range. Zoologicheskii zhurnal, 2000, vol. 79, iss. 9, pp. 1133–1136 (in Russian).Baskevich M. I., Oparin M. L. On new finding of the birch mouse Sicista strandi (Rodentia, Dipodoidea) specifying the northeastern boundary of the species range. Zoologicheskii zhurnal, 2000, vol. 79, iss. 9, pp. 1133–1136 (in Russian).Baskevich M. I., Oparin M. L. Multiple chromosomal polymorphism in Birch mice of the subtilis group (Rodentia, Dipodoidea, Sicista) from the Saratov right bank of the Volga river. Biology Bulletin, 2023, vol. 50, iss. 3, pp. 300–310. https://doi.org/10.1134/s1062359022700066Baskevich M. I., Oparin M. L. Multiple chromosomal polymorphism in Birch mice of the subtilis group (Rodentia, Dipodoidea, Sicista) from the Saratov right bank of the Volga river. Biology Bulletin, 2023, vol. 50, iss. 3, pp. 300–310. https://doi.org/10.1134/s1062359022700066Baskevich M. I., Oparin M. L., Illarionova N. A. Cytogenetic and molecular genetic differentiation in the group of steppe voles (Rodentia, Dipodoidea, Sicista). In: Averianov A. O., Abramson N. I., eds. Systematics, Phylogeny, and Paleontology of Small Mammals. Proceedings of the International Conference devoted to the 90-th anniversary of Prof. I. M. Gromov. St. Petersburg, Zoological Institute of the Russian Academy of Sciences Publ., 2003, pp. 44–46 (in Russian).Baskevich M. I., Oparin M. L., Illarionova N. A. Cytogenetic and molecular genetic differentiation in the group of steppe voles (Rodentia, Dipodoidea, Sicista). In: Averianov A. O., Abramson N. I., eds. Systematics, Phylogeny, and Paleontology of Small Mammals. Proceedings of the International Conference devoted to the 90-th anniversary of Prof. I. M. Gromov. St. Petersburg, Zoological Institute of the Russian Academy of Sciences Publ., 2003, pp. 44–46 (in Russian).Baskevich M. I., Okulova N. M., Potapov S. G., Illarionova N. A., Krysanov E. Yu., Shchipanov N. A., Oparin M. L., Vlasov A. A. On the diagnostics and distribution of sibling species of Sicista (Rodentia, Dipodoidea) on the territory of Russian plain and Caucasus. Proceedings of the Zoological Institute RAS, 2005, vol. 306, pp. 22–40 (in Russian).Baskevich M. I., Okulova N. M., Potapov S. G., Illarionova N. A., Krysanov E. Yu., Shchipanov N. A., Oparin M. L., Vlasov A. A. On the diagnostics and distribution of sibling species of Sicista (Rodentia, Dipodoidea) on the territory of Russian plain and Caucasus. Proceedings of the Zoological Institute RAS, 2005, vol. 306, pp. 22–40 (in Russian).Baskevich M. I., Oparin M. L., Cherepanova E. V., Avilova E. A. Chromosome differentiation of Sicista subtilis (Rodentia, Dipodoidea) in the Low Volga river basin. Zoologicheskii zhurnal, 2010, vol. 89, iss. 6, pp. 749–757 (in Russian).Baskevich M. I., Oparin M. L., Cherepanova E. V., Avilova E. A. Chromosome differentiation of Sicista subtilis (Rodentia, Dipodoidea) in the Low Volga river basin. Zoologicheskii zhurnal, 2010, vol. 89, iss. 6, pp. 749–757 (in Russian).Baskevich M. I., Potapov S. G., Mironova T. A. Caucasian cryptic species of rodents as models in research on the problems of species and speciation. Biology Bulletin Reviews, 2016, vol. 6, iss. 3, pp. 245–259. https://doi.org/10.1134/S2079086416030026Baskevich M. I., Potapov S. G., Mironova T. A. Caucasian cryptic species of rodents as models in research on the problems of species and speciation. Biology Bulletin Reviews, 2016, vol. 6, iss. 3, pp. 245–259. https://doi.org/10.1134/S2079086416030026Baskevich M. I., Bogdanov A. S., Khlyap L. A. Taxonomy and phylogeny of sibling-species Sicista of the group caucasica and their position in the genus Sicista (Rodentia, Dipodoidea) according to sequencing of the IRBP gene fragment of nuclear DNA. Biology Bulletin, 2018, vol. 45, iss. 5, pp. 432–437. https://doi.org/10.1134/S1062359018050047Baskevich M. I., Bogdanov A. S., Khlyap L. A. Taxonomy and phylogeny of sibling-species Sicista of the group caucasica and their position in the genus Sicista (Rodentia, Dipodoidea) according to sequencing of the IRBP gene fragment of nuclear DNA. Biology Bulletin, 2018, vol. 45, iss. 5, pp. 432–437. https://doi.org/10.1134/S1062359018050047Baskevich M. I., Bogdanov A. S., Khlyap L. A., Malygin V. M., Oparin M. L., Sapelnikov S. F., Sheftel B. I. Phylogeny and differentiation of sibling-species Sicista of the group Betulina (Rodentia, Dipodoidea): Results of analysis of a fragment of the IRBP gene of nuclear DNA variability. Biology Bulletin, 2020, vol. 47, iss. 5, pp. 482–489. https://doi.org/10.1134/S1062359018050047Baskevich M. I., Bogdanov A. S., Khlyap L. A., Malygin V. M., Oparin M. L., Sapelnikov S. F., Sheftel B. I. Phylogeny and differentiation of sibling-species Sicista of the group Betulina (Rodentia, Dipodoidea): Results of analysis of a fragment of the IRBP gene of nuclear DNA variability. Biology Bulletin, 2020, vol. 47, iss. 5, pp. 482–489. https://doi.org/10.1134/S1062359018050047Baskevich M. I., Khlyap L. A., Bogdanov A. S. Strand’s birch mouse Sicista strandi (Rodentia, Dipodoidea) on the southwestern periphery of the species range: Genetic and ecological aspects. Biology Bulletin, 2024, vol. 51, iss. 10, pp. 3031–3040. https://doi.org/10.1134/S1062359024700791Baskevich M. I., Khlyap L. A., Bogdanov A. S. Strand’s birch mouse Sicista strandi (Rodentia, Dipodoidea) on the southwestern periphery of the species range: Genetic and ecological aspects. Biology Bulletin, 2024, vol. 51, iss. 10, pp. 3031–3040. https://doi.org/10.1134/S1062359024700791Bogdanov A. S., Rozhkova D. N., Khlyap L. A., Baskevich M. I. Multi-level molecular differentiation of populations of the Strand’s birch mouse Sicista strandi (Rodentia, Dipodoidea). Animals, 2025, vol. 15, art. 2605. https://doi.org/10.3390/ani15172605Bogdanov A. S., Rozhkova D. N., Khlyap L. A., Baskevich M. I. Multi-level molecular differentiation of populations of the Strand’s birch mouse Sicista strandi (Rodentia, Dipodoidea). Animals, 2025, vol. 15, art. 2605. https://doi.org/10.3390/ani15172605Bystrakova N. V. Taxonomic and Genetic Diversity of Small Mammals of the Middle Volga Region. Thesis Diss. Cand. Sci. (Biol.). Moscow, 2000. 24 p. (in Russian).Bystrakova N. V. Taxonomic and Genetic Diversity of Small Mammals of the Middle Volga Region. Thesis Diss. Cand. Sci. (Biol.). Moscow, 2000. 24 p. (in Russian).Bystrakova N. V., Ermakov O. A. Southern birch mouse Sicista subtilis Pallas, 1773. In: Lissovsky A. A., Stakheev V. V., Saveljev A. P., Ermakov O. A., Smirnov D. G., Glazov D. M., Obolenskaya E. V., Sheftel B. I., Titov S. V., eds. Atlas of Mammal Distribution in the European Part of Russia. Moscow, KMK Scientific Press, 2025, pp. 300 (in Russian).Bystrakova N. V., Ermakov O. A. Southern birch mouse Sicista subtilis Pallas, 1773. In: Lissovsky A. A., Stakheev V. V., Saveljev A. P., Ermakov O. A., Smirnov D. G., Glazov D. M., Obolenskaya E. V., Sheftel B. I., Titov S. V., eds. Atlas of Mammal Distribution in the European Part of Russia. Moscow, KMK Scientific Press, 2025, pp. 300 (in Russian).Cserkész T., Aczél-Fridrich Z., Hegyeli Z., Sugár S., Czabán D., Horváth O., Sramkó G. Rediscovery of the Hungarian birch mouse (Sicista subtilis trizona) in Transylvania (Romania) with molecular characterisation of its phylogenetic affinities. Mammalia, 2015, vol. 79, iss. 2, pp. 215–224. https://doi.org/10.1515/mammalia-2013-0167Cserkész T., Aczél-Fridrich Z., Hegyeli Z., Sugár S., Czabán D., Horváth O., Sramkó G. Rediscovery of the Hungarian birch mouse (Sicista subtilis trizona) in Transylvania (Romania) with molecular characterisation of its phylogenetic affinities. Mammalia, 2015, vol. 79, iss. 2, pp. 215–224. https://doi.org/10.1515/mammalia-2013-0167Cserkész T., Rusin M., Sramko G. An integrative systematic revision of the European southern birch mice (Rodentia: Sminthidae, Sicista subtilis group). Mammal Review, 2016, vol. 46, iss. 2, pp. 114–130. https://doi.org/10.1111/mam.12058Cserkész T., Rusin M., Sramko G. An integrative systematic revision of the European southern birch mice (Rodentia: Sminthidae, Sicista subtilis group). Mammal Review, 2016, vol. 46, iss. 2, pp. 114–130. https://doi.org/10.1111/mam.12058Cserkész T., Fulop A., Almerekova Sh., Kondor T., Levente L., Sramko G. Phylogenetic and morphological analysis of birch mice (genus Sicista, family Smintidae, Rodentia) in the Kazak Gradle with description of a new species. Journal of Mammalian Evolution, 2019, vol. 26, iss. 1, pp. 147–161. https://doi.org/10.1007/s10914-017-9409-6Cserkész T., Fulop A., Almerekova Sh., Kondor T., Levente L., Sramko G. Phylogenetic and morphological analysis of birch mice (genus Sicista, family Smintidae, Rodentia) in the Kazak Gradle with description of a new species. Journal of Mammalian Evolution, 2019, vol. 26, iss. 1, pp. 147–161. https://doi.org/10.1007/s10914-017-9409-6Deng Yu., Liu Q., Wang X., Li B. V., Wang J., Liu Sh., Liao R., Liu Sh., Chen Sh. Molecular phylogeny and taxonomy of the Sicista concolor group (Mammalia, Rodentia, Sicistidae) with the description of a new species. Zoosystematics and Evolution, 2025, vol. 101, iss. 3, pp. 1197–1211. https://doi.org/10.3897/zse.101.155510Deng Yu., Liu Q., Wang X., Li B. V., Wang J., Liu Sh., Liao R., Liu Sh., Chen Sh. Molecular phylogeny and taxonomy of the Sicista concolor group (Mammalia, Rodentia, Sicistidae) with the description of a new species. Zoosystematics and Evolution, 2025, vol. 101, iss. 3, pp. 1197–1211. https://doi.org/10.3897/zse.101.155510Ellerman J. R., Morrison-Scott T. C. S. Checklist of the Palearctic and Indian Mammals, 1758 to 1946. London, British Museum of Natural History, 1951. 810 p.Ellerman J. R., Morrison-Scott T. C. S. Checklist of the Palearctic and Indian Mammals, 1758 to 1946. London, British Museum of Natural History, 1951. 810 p.Emelyanova A. A., Sidorova O. V. Comparison of micromammalian faunas of natural and anthropogenic biotopes of Darwin and Central Forest Reservs over the period 2005–2009. Bulletin of Tver State University. Series: Biology and Ecology, 2014, iss. 2, pp. 48–62 (in Russian).Emelyanova A. A., Sidorova O. V. Comparison of micromammalian faunas of natural and anthropogenic biotopes of Darwin and Central Forest Reservs over the period 2005–2009. Bulletin of Tver State University. Series: Biology and Ecology, 2014, iss. 2, pp. 48–62 (in Russian).Ford C. E., Hamerton J. L. A colchicine hypotonic citrate, squash sequence for mammalian chromosomes. Stain Technology, 1956, vol. 31, iss. 6, pp. 247–251. https://doi.org/10.3109/10520295609113814Ford C. E., Hamerton J. L. A colchicine hypotonic citrate, squash sequence for mammalian chromosomes. Stain Technology, 1956, vol. 31, iss. 6, pp. 247–251. https://doi.org/10.3109/10520295609113814Holden M. E., Musser G. G. Superfamily Dipodoidea. In: Wilson D. E., Reeder D. M., eds. Mammal Species of the World. Baltimore, Johns Hopkins University Press, 2005, pp. 871–893.Holden M. E., Musser G. G. Superfamily Dipodoidea. In: Wilson D. E., Reeder D. M., eds. Mammal Species of the World. Baltimore, Johns Hopkins University Press, 2005, pp. 871–893.Holden M. E., Cserkész T., Musser G. Family Sminthidae. In: Wilson D. E., Lacher T. E., Jr., Mittermeier R. A., eds. Handbook of the Mammals of the World. Barcelona, Lynx Ediciones, 2017, vol. 7, pp. 22–48.Holden M. E., Cserkész T., Musser G. Family Sminthidae. In: Wilson D. E., Lacher T. E., Jr., Mittermeier R. A., eds. Handbook of the Mammals of the World. Barcelona, Lynx Ediciones, 2017, vol. 7, pp. 22–48.Karpenko N. T. Characteristics of the species composition of mammals, the problem of preservation of rare and endangered species “Bogdinsko-Baskunchak” reserve. Samarskaya Luka: Problems of Regional and Global Ecology, 2013, vol. 22, no. 4, pp. 122–127 (in Russian).Karpenko N. T. Characteristics of the species composition of mammals, the problem of preservation of rare and endangered species “Bogdinsko-Baskunchak” reserve. Samarskaya Luka: Problems of Regional and Global Ecology, 2013, vol. 22, no. 4, pp. 122–127 (in Russian).Kovalskaya Yu. M., Fedorovich E. Yu. To distributing chromosome forms of the Southern birch mouse Sicista subtilis (Rodentia, Dipodoidea). Zoologicheskii zhurnal, 1997, vol. 76, iss. 12, pp. 1430–1433 (in Russian).Kovalskaya Yu. M., Fedorovich E. Yu. To distributing chromosome forms of the Southern birch mouse Sicista subtilis (Rodentia, Dipodoidea). Zoologicheskii zhurnal, 1997, vol. 76, iss. 12, pp. 1430–1433 (in Russian).Kovalskaya Yu. M., Tikhonov I. A., Tikhonova G. N., Surov A. V., Bogomolov P. L. New geographical localities of chromosome forms of southern birch mouse (subtilis group) and description of Sicista severtzovi cimlanica subsp. n. (Mammalia, Rodentia) from the Middle Don river basin. Zoologicheskii zhurnal, 2000, vol. 79, iss. 8, pp. 954–964 (in Russian).Kovalskaya Yu. M., Tikhonov I. A., Tikhonova G. N., Surov A. V., Bogomolov P. L. New geographical localities of chromosome forms of southern birch mouse (subtilis group) and description of Sicista severtzovi cimlanica subsp. n. (Mammalia, Rodentia) from the Middle Don river basin. Zoologicheskii zhurnal, 2000, vol. 79, iss. 8, pp. 954–964 (in Russian).Kovalskaya Yu. M., Aniskin V. M., Bogomolov P. L., Surov A. V., Tikhonov I. A., Tikhonova G. N., Robinson T. J., Volobouev V. T. Karyotype reorganisation in the subtilis group of birch mice (Rodentia, Dipodidae, Sicista): Unexpected taxonomic diversity within a limited distribution. Cytogenetic and Genome Research, 2011, vol. 132, iss. 4, pp. 271–288. https://doi.org/10.1159/000322823Kovalskaya Yu. M., Aniskin V. M., Bogomolov P. L., Surov A. V., Tikhonov I. A., Tikhonova G. N., Robinson T. J., Volobouev V. T. Karyotype reorganisation in the subtilis group of birch mice (Rodentia, Dipodidae, Sicista): Unexpected taxonomic diversity within a limited distribution. Cytogenetic and Genome Research, 2011, vol. 132, iss. 4, pp. 271–288. https://doi.org/10.1159/000322823Kubantsev B. S., Uvarova V. Ya., Kosareva N. A. The Fauna of the Volgograd Region. In: Nazemnye pozvonochnye zhivotnye [Terrestrial Vertebrates]. Volgograd, Volgogradskoe knizhnoe izdatel'stvo, 1962, pp. 51–155 (in Russian).Kubantsev B. S., Uvarova V. Ya., Kosareva N. A. The Fauna of the Volgograd Region. In: Nazemnye pozvonochnye zhivotnye [Terrestrial Vertebrates]. Volgograd, Volgogradskoe knizhnoe izdatel'stvo, 1962, pp. 51–155 (in Russian).Kurochkin D. V., Korshunov E. N., Ponimatko A. O. An Annotated List of Mammals in the Kerzhensky Nature Reserve. Proceedings of the Kerzhensky State Nature Reserve, 2002, vol. 2, pp. 9–23 (in Russian).Kurochkin D. V., Korshunov E. N., Ponimatko A. O. An Annotated List of Mammals in the Kerzhensky Nature Reserve. Proceedings of the Kerzhensky State Nature Reserve, 2002, vol. 2, pp. 9–23 (in Russian).Lebedev V., Poplavskaya N., Bannikova A., Rusin M., Surov A., Kovalskaya Yu. Genetic variation in the Sicista subtilis (Pallas, 1773) species group (Rodentia, Sminthidae), as compared to karyotype differentiation. Mammalia, 2019a, vol. 84, iss. 2, pp. 185–194. https://doi.org/10.1515/mammalia-2018-0216Lebedev V., Poplavskaya N., Bannikova A., Rusin M., Surov A., Kovalskaya Yu. Genetic variation in the Sicista subtilis (Pallas, 1773) species group (Rodentia, Sminthidae), as compared to karyotype differentiation. Mammalia, 2019a, vol. 84, iss. 2, pp. 185–194. https://doi.org/10.1515/mammalia-2018-0216Lebedev V. S., Rusin M. Yu., Zemlemerova E. D., MatrosovaV. A., Bannikova A. A., Kovalskaya Yu. M., Tesakov A. S. Phylogeny and evolutionary history of birch mice Sicista Griffith, 1827 (Sminthidae, Rodentia): Implications from a multigene study. Journal of Zoological Systematics and Evolutionary Research, 2019b, vol. 57, iss. 3, pp. 695–709. https://doi.org/10.1111/jzs.12279Lebedev V. S., Rusin M. Yu., Zemlemerova E. D., MatrosovaV. A., Bannikova A. A., Kovalskaya Yu. M., Tesakov A. S. Phylogeny and evolutionary history of birch mice Sicista Griffith, 1827 (Sminthidae, Rodentia): Implications from a multigene study. Journal of Zoological Systematics and Evolutionary Research, 2019b, vol. 57, iss. 3, pp. 695–709. https://doi.org/10.1111/jzs.12279Lebedev V. S., Kovalskaya Yu., Solovyeva E. N., Zemlemerova E. D., Bannikov A. A., Rusin M. Yu., Matrosova V. A. Molecular systematics of the Sicista tianschanica species complex: A contribution from historical DNA analysis. PeerJ, 2021, vol. 9, art. e10759. https://doi.org/10.7717/peerj.10759Lebedev V. S., Kovalskaya Yu., Solovyeva E. N., Zemlemerova E. D., Bannikov A. A., Rusin M. Yu., Matrosova V. A. Molecular systematics of the Sicista tianschanica species complex: A contribution from historical DNA analysis. PeerJ, 2021, vol. 9, art. e10759. https://doi.org/10.7717/peerj.10759Moskvitchev A. N. A Brief systematic list of Mammals in the Ulyanovsk region. Priroda Simbirskogo Povolzh'ya [Nature of the Simbirsk Volga Region]. Ulyanovsk, I. N. Ulyanov Ulyanovsk State Pedagogical University Publ., 2022, iss. 23, pp. 99–109 (in Russian).Moskvitchev A. N. A Brief systematic list of Mammals in the Ulyanovsk region. Priroda Simbirskogo Povolzh'ya [Nature of the Simbirsk Volga Region]. Ulyanovsk, I. N. Ulyanov Ulyanovsk State Pedagogical University Publ., 2022, iss. 23, pp. 99–109 (in Russian).Ognev S. I. Zveri SSSR i prilezhashchikh stran. Gryzuny [Mammals of USSR and Adjacent Countries: Rodents]. Moscow, Izdatel'stvo AN SSSR, 1948, vol. 6. 559 p. (in Russian).Ognev S. I. Zveri SSSR i prilezhashchikh stran. Gryzuny [Mammals of USSR and Adjacent Countries: Rodents]. Moscow, Izdatel'stvo AN SSSR, 1948, vol. 6. 559 p. (in Russian).Ogureeva G. N., Leonova N. B., Miklyaeva I. M., Bocharnikov M. V., Fedosov V. E., Muchnik E. E., Urbanavichus G. P., Emelyanova L. G., Khlyap L. A., Rumyantsev V. Yu., Kuzikov I. V., Lipka O. N., Arkhipova M. V., Buldakova E. V., Kadetov N. G. The Biodiversity of Russian Biomes. The Biomes of Plains. Moscow, Institute of Global Climate and Ecology Publ., 2020. 623 p. (in Russian).Ogureeva G. N., Leonova N. B., Miklyaeva I. M., Bocharnikov M. V., Fedosov V. E., Muchnik E. E., Urbanavichus G. P., Emelyanova L. G., Khlyap L. A., Rumyantsev V. Yu., Kuzikov I. V., Lipka O. N., Arkhipova M. V., Buldakova E. V., Kadetov N. G. The Biodiversity of Russian Biomes. The Biomes of Plains. Moscow, Institute of Global Climate and Ecology Publ., 2020. 623 p. (in Russian).Okulova N. M., Khelevina S. A. Melkie lesnye mlekopitayushchie Ivanovskoi oblasti i ee okrestnostei [Small Forest Mammals of the Ivanovo Region and Its Surroundings]. Ivanovo, Ivanovo State University Publ., 1989. 100 p. (in Russian).Okulova N. M., Khelevina S. A. Melkie lesnye mlekopitayushchie Ivanovskoi oblasti i ee okrestnostei [Small Forest Mammals of the Ivanovo Region and Its Surroundings]. Ivanovo, Ivanovo State University Publ., 1989. 100 p. (in Russian).Oparin M. L., Tikhonov I. A., Kovalskaya Yu. M., Bogomolov P. L., Shapovalov A. S. To the distribution of the dark birch mouse Sicista severtzovi Ognev, 1935 (Mammalia) on the Russian Plain. In: Gavrilov V. M., ed. The Role of Biological Stations in the Conservation of Russian: Proceedings of the conference, dedicated to 250th anniversary of Lomonosov Moscow State University and the 90th anniversary of the S. N. Skadovsky Zvenigorod Biological Station. Moscow, Oikos, 2001, pp. 121–123 (in Russian).Oparin M. L., Tikhonov I. A., Kovalskaya Yu. M., Bogomolov P. L., Shapovalov A. S. To the distribution of the dark birch mouse Sicista severtzovi Ognev, 1935 (Mammalia) on the Russian Plain. In: Gavrilov V. M., ed. The Role of Biological Stations in the Conservation of Russian: Proceedings of the conference, dedicated to 250th anniversary of Lomonosov Moscow State University and the 90th anniversary of the S. N. Skadovsky Zvenigorod Biological Station. Moscow, Oikos, 2001, pp. 121–123 (in Russian).Order No. 75 of the Ministry of Natural Resources and Environmental Protection of the Republic of Kalmykia, dated May 20, 2025, “On the Approval of Lists of Fauna and Flora Species Included in the Red Book of the Republic of Kalmykia”. Elista, Ministry of Natural Resources and Environmental Protection of the Republic of Kalmykia Publ., 2025. 15 p. Available at: https://kalmpriroda.ru (accessed July 24, 2025) (in Russian).Order No. 75 of the Ministry of Natural Resources and Environmental Protection of the Republic of Kalmykia, dated May 20, 2025, “On the Approval of Lists of Fauna and Flora Species Included in the Red Book of the Republic of Kalmykia”. Elista, Ministry of Natural Resources and Environmental Protection of the Republic of Kalmykia Publ., 2025. 15 p. Available at: https://kalmpriroda.ru (accessed July 24, 2025) (in Russian).Popov V. A. Mlekopitayushchie Volzhsko-Kamskogo kraya: nasekomoyadnye, rukokrylye, gryzuny [Mammals of the Volga-Kama Region: Insectivores, Bats, and Rodents]. Kazan, Kazan Branch of the Academy of Sciences of the USSR Publ., 1960. 134 p. (in Russian).Popov V. A. Mlekopitayushchie Volzhsko-Kamskogo kraya: nasekomoyadnye, rukokrylye, gryzuny [Mammals of the Volga-Kama Region: Insectivores, Bats, and Rodents]. Kazan, Kazan Branch of the Academy of Sciences of the USSR Publ., 1960. 134 p. (in Russian).Potapov S. K., Bugaev K. E., Artaev O. N. Mammals. Flora and Fauna of Nature Reserves. Iss. 120. Vertebrates of the Mordovian Nature Reserve. Moscow, Russian Academy of Sciences Commission on the Conservation of Biological Diversity, 2012, pp. 50–61 (in Russian).Potapov S. K., Bugaev K. E., Artaev O. N. Mammals. Flora and Fauna of Nature Reserves. Iss. 120. Vertebrates of the Mordovian Nature Reserve. Moscow, Russian Academy of Sciences Commission on the Conservation of Biological Diversity, 2012, pp. 50–61 (in Russian).Priroda Kostromskoi oblasti i ee okhrana [The Nature of the Kostroma Region and its Conservation]. Yaroslavl, Verkhne-Volzhskoe knizhnoe izdatel'stvo, 1987. 78 p. (in Russian).Priroda Kostromskoi oblasti i ee okhrana [The Nature of the Kostroma Region and its Conservation]. Yaroslavl, Verkhne-Volzhskoe knizhnoe izdatel'stvo, 1987. 78 p. (in Russian).Red Book of Astrakhan Region. Third edition. Astrakhan, Nature Management and Environmental Protection Service of the Astrakhan Region Publ., 2024. 615 p. (in Russian).Red Book of Astrakhan Region. Third edition. Astrakhan, Nature Management and Environmental Protection Service of the Astrakhan Region Publ., 2024. 615 p. (in Russian).Red Book of the City of Moscow. 3rd ed., rev. and expanded. Moscow, Department of Natural Resource Management and Environmental Protection of the City of Moscow, 2022. 848 p. (in Russian).Red Book of the City of Moscow. 3rd ed., rev. and expanded. Moscow, Department of Natural Resource Management and Environmental Protection of the City of Moscow, 2022. 848 p. (in Russian).Red Book of the Kostroma Region. Kostroma, Kostroma State University Publ., 2009. 367 p. (in Russian).Red Book of the Kostroma Region. Kostroma, Kostroma State University Publ., 2009. 367 p. (in Russian).Red Book of the Nizhni Novgorod Region. Vol. 1. Animals. 3rd ed., rev. and expanded. Nizhni Novgorod, Ecotsenter “Dront”, 2026. 444 p. (in Russian).Red Book of the Nizhni Novgorod Region. Vol. 1. Animals. 3rd ed., rev. and expanded. Nizhni Novgorod, Ecotsenter “Dront”, 2026. 444 p. (in Russian).Red Book of the Penza Region. Vol. 2. Animals. 2nd ed. Voronezh, Izdatel'stvo im. E. A. Bolkhovitinova, 2019. 264 p. (in Russian).Red Book of the Penza Region. Vol. 2. Animals. 2nd ed. Voronezh, Izdatel'stvo im. E. A. Bolkhovitinova, 2019. 264 p. (in Russian).Red Book of the Republic of Kalmykia: in 2 vols. Vol. 1: Rare and Endangered Animals. Elista, Dzhangar, 2013. 200 p.Red Book of the Republic of Kalmykia: in 2 vols. Vol. 1: Rare and Endangered Animals. Elista, Dzhangar, 2013. 200 p.Red Book of the Republic of Mordovia. Vol. 2. Animals. Saransk, Mordovskoe knizhnoe izdatel'stvo, 2005. 336 p. (in Russian).Red Book of the Republic of Mordovia. Vol. 2. Animals. Saransk, Mordovskoe knizhnoe izdatel'stvo, 2005. 336 p. (in Russian).Red Book of the Republic of Tatarstan (Animals, Plants, Fungi). Fourth edition. Kazan, Academy of Sciences of the Republic of Tatarstan Publ., 2025. 804 p. (in Russian).Red Book of the Republic of Tatarstan (Animals, Plants, Fungi). Fourth edition. Kazan, Academy of Sciences of the Republic of Tatarstan Publ., 2025. 804 p. (in Russian).Red Book of the Ulyanovsk Region. Ulyanovsk, Buki Vedi, 2015. 550 p. (in Russian).Red Book of the Ulyanovsk Region. Ulyanovsk, Buki Vedi, 2015. 550 p. (in Russian).Red Book of the Ulyanovsk Region. Ulyanovsk, “Povolzhye” Research Center, 2025. 670 p. (in Russian).Red Book of the Ulyanovsk Region. Ulyanovsk, “Povolzhye” Research Center, 2025. 670 p. (in Russian).Red Book of the Yaroslavl Region. Yaroslavl, Akademiya 76, 2015. 470 p. (in Russian).Red Book of the Yaroslavl Region. Yaroslavl, Akademiya 76, 2015. 470 p. (in Russian).Red Data Book of Russian Federation. Animals. 2nd ed. Moscow, VNII Ecology Publ., 2021. 1128 p. (in Russian).Red Data Book of Russian Federation. Animals. 2nd ed. Moscow, VNII Ecology Publ., 2021. 1128 p. (in Russian).Resolutions of the Government of the Yaroslavl Region No. 239-p dated March 6, 2015, and No. 1200-p dated November 9, 2015, 2015 “On the Approval of Lists of Species of Fungi, Lichens, Plants, and Animals Included in the Red Book of the Yaroslavl Region and Excluded from the Red Book of the Yaroslavl Region”. Yaroslavl, 2015. 31 p. Available at: https://www.yarregion.ru (accessed on July 24, 2025) (in Russian).Resolutions of the Government of the Yaroslavl Region No. 239-p dated March 6, 2015, and No. 1200-p dated November 9, 2015, 2015 “On the Approval of Lists of Species of Fungi, Lichens, Plants, and Animals Included in the Red Book of the Yaroslavl Region and Excluded from the Red Book of the Yaroslavl Region”. Yaroslavl, 2015. 31 p. Available at: https://www.yarregion.ru (accessed on July 24, 2025) (in Russian).Rigina Е. Yu. The fauna of the rodent species (Rodentia) in the Samara region. In: Chibilev A. A., ed. Steppes of Northern Eurasia: Materials of the IV International Symposium. Orenburg, IPK “Gazprompechat”, 2006, pp. 602–605 (in Russian).Rigina Е. Yu. The fauna of the rodent species (Rodentia) in the Samara region. In: Chibilev A. A., ed. Steppes of Northern Eurasia: Materials of the IV International Symposium. Orenburg, IPK “Gazprompechat”, 2006, pp. 602–605 (in Russian).Rusin M., Lebedev V., Matrosova V., Zemlemerova E., Lopatina N., Bannikova A. Hidden diversity in the Caucasian mountains: An example of birch mice (Rodentia, Sminthidae, Sicista). Hystrix, 2018, vol. 29, iss. 1, pp. 61–66. https://doi.org/10.4404/hystrix-00050-2018Rusin M., Lebedev V., Matrosova V., Zemlemerova E., Lopatina N., Bannikova A. Hidden diversity in the Caucasian mountains: An example of birch mice (Rodentia, Sminthidae, Sicista). Hystrix, 2018, vol. 29, iss. 1, pp. 61–66. https://doi.org/10.4404/hystrix-00050-2018Sambrook J., Fritsch E. F., Maniatis T. Molecular Cloning: A Laboratory Manual. New York, Cold Spring Harbor Laboratory Press, 1989. 398 p.Sambrook J., Fritsch E. F., Maniatis T. Molecular Cloning: A Laboratory Manual. New York, Cold Spring Harbor Laboratory Press, 1989. 398 p.Sapozhenkov Yu. F. New mammal findings in the Kostroma region. In: Zhivotnyi mir Kostromskoi oblasti [The Fauna of the Kostroma Region]. Kostroma, Kostroma State Pedagogical Institute named after N. A. Nekrasov Publ., 1973, iss. 31, pp. 7–10 (in Russian).Sapozhenkov Yu. F. New mammal findings in the Kostroma region. In: Zhivotnyi mir Kostromskoi oblasti [The Fauna of the Kostroma Region]. Kostroma, Kostroma State Pedagogical Institute named after N. A. Nekrasov Publ., 1973, iss. 31, pp. 7–10 (in Russian).Shemyatikhina G. B., Nafeev A. A., Krivosheyev V. A. Specific compound and biotopical coordinates of small mammals from entomophagous and rodents group in territory of Ulyanovsk oblast. Proceedings of the Samara Scientific Center of the Russian Academy of Sciences, 2009, vol. 11, no. 1, pp. 184–187 (in Russian).Shemyatikhina G. B., Nafeev A. A., Krivosheyev V. A. Specific compound and biotopical coordinates of small mammals from entomophagous and rodents group in territory of Ulyanovsk oblast. Proceedings of the Samara Scientific Center of the Russian Academy of Sciences, 2009, vol. 11, no. 1, pp. 184–187 (in Russian).Shenbrot G. I., Sokolov V. E., Geptner V. G., Kovalskaya Yu. M. Dipodoidae. Mammals of Russia Andadjacent Regions. Moscow, Nauka, 1995. 573 p. (in Russian).Shenbrot G. I., Sokolov V. E., Geptner V. G., Kovalskaya Yu. M. Dipodoidae. Mammals of Russia Andadjacent Regions. Moscow, Nauka, 1995. 573 p. (in Russian).Shlyakhtin G. V., Il'in V. Yu., Oparin M. L., Belyachenko A. V., Bystrakova N. V., Ermakov O. A., Zavialov E. V., Zakharov K. S., Kaibeleva E. I., Koshkin V. A., Kurmaeva N. M., Lukyanov S. B., Mosolova E. Yu., Oparina O. S., Semikhatova S. N., Smirnov D. G., Sonin K. A., Tabachishin V. G., Titov S. V., Filippiev A. O., Khuchraev S. O., Yakushev N. N. Mammals of the Northern Lower-Volga region: In 3 vols. Vol. I. Theriofauna composition. Saratov, Saratov University Press, 2009. 248 p. (in Russian).Shlyakhtin G. V., Il'in V. Yu., Oparin M. L., Belyachenko A. V., Bystrakova N. V., Ermakov O. A., Zavialov E. V., Zakharov K. S., Kaibeleva E. I., Koshkin V. A., Kurmaeva N. M., Lukyanov S. B., Mosolova E. Yu., Oparina O. S., Semikhatova S. N., Smirnov D. G., Sonin K. A., Tabachishin V. G., Titov S. V., Filippiev A. O., Khuchraev S. O., Yakushev N. N. Mammals of the Northern Lower-Volga region: In 3 vols. Vol. I. Theriofauna composition. Saratov, Saratov University Press, 2009. 248 p. (in Russian).Sidorova O. V. Specific variety and biotope distribution fine mammal in natural and anthropogenic changed biotope Darvinskogo and Central-Forest State Reserve. Bulletin of Tver State University. Series: Biology and Ecology, 2009, iss. 14, pp. 100–109 (in Russian).Sidorova O. V. Specific variety and biotope distribution fine mammal in natural and anthropogenic changed biotope Darvinskogo and Central-Forest State Reserve. Bulletin of Tver State University. Series: Biology and Ecology, 2009, iss. 14, pp. 100–109 (in Russian).Sokolov V. E., Baskevich M. I. A new species of unstriped birch mice (Rodentia, Dipodoidea) from the Minor Caucasus. Zoologicheskii zhurnal, 1988, vol. 67, iss. 2, pp. 300–304 (in Russian).Sokolov V. E., Baskevich M. I. A new species of unstriped birch mice (Rodentia, Dipodoidea) from the Minor Caucasus. Zoologicheskii zhurnal, 1988, vol. 67, iss. 2, pp. 300–304 (in Russian).Sokolov V. E., Kovalskaya Yu. M. The system of the genus Sicista and chromosomal forms of Tien Shan birch mice, S. tianschanica Salensky, 1903. In: Abstracts of Papers of the V Congress of the All-Union Theriological Society. Moscow, USSR Academy of Sciences Publ., 1990a, pt. 1, pp. 99–100 (in Russian).Sokolov V. E., Kovalskaya Yu. M. The system of the genus Sicista and chromosomal forms of Tien Shan birch mice, S. tianschanica Salensky, 1903. In: Abstracts of Papers of the V Congress of the All-Union Theriological Society. Moscow, USSR Academy of Sciences Publ., 1990a, pt. 1, pp. 99–100 (in Russian).Sokolov V. E., Kovalskaya Yu. M. Zoologicheskii zhurnal, 1990b, vol. 69, iss. 9, pp. 152–157 (in Russian).Sokolov V. E., Kovalskaya Yu. M. Zoologicheskii zhurnal, 1990b, vol. 69, iss. 9, pp. 152–157 (in Russian).Sokolov V. E., Baskevich M. I., Kovalskaya, Yu. M., Revision of unstriped birch mice of the Caucasus: Sibling species Sicista caucasica Vinigradov, 1925 and S. kluchorica sp. n. (Rodentia, Dipodidae). Zoologicheskii zhurnal, 1981, vol. 60, iss. 9, pp. 1386–1393 (in Russian).Sokolov V. E., Baskevich M. I., Kovalskaya, Yu. M., Revision of unstriped birch mice of the Caucasus: Sibling species Sicista caucasica Vinigradov, 1925 and S. kluchorica sp. n. (Rodentia, Dipodidae). Zoologicheskii zhurnal, 1981, vol. 60, iss. 9, pp. 1386–1393 (in Russian).Sokolov V. E., Baskevich M. I., Kovalskaya Yu. M. Sicista kazbegica sp. n. (Rodentia, Dipodidae) from the basin of the upper reaches of the Terek river. Zoologicheskii zhurnal, 1986a, vol. 65, iss. 6, pp. 949–952 (in Russian).Sokolov V. E., Baskevich M. I., Kovalskaya Yu. M. Sicista kazbegica sp. n. (Rodentia, Dipodidae) from the basin of the upper reaches of the Terek river. Zoologicheskii zhurnal, 1986a, vol. 65, iss. 6, pp. 949–952 (in Russian).Sokolov V. E., Baskevich M. I., Kovalskaya Yu. M., Variability of the karyotype of the steppe birch mouse Sicista subtilis Pallas (1778) and justification of the species independence of S. severtzovi Ognev, 1935 (Rodentia, Zapodidae). Zoologicheskii zhurnal, 1986b, vol. 65, iss. 8, pp. 1684–1692 (in Russian).Sokolov V. E., Baskevich M. I., Kovalskaya Yu. M., Variability of the karyotype of the steppe birch mouse Sicista subtilis Pallas (1778) and justification of the species independence of S. severtzovi Ognev, 1935 (Rodentia, Zapodidae). Zoologicheskii zhurnal, 1986b, vol. 65, iss. 8, pp. 1684–1692 (in Russian).Sokolov V. E., Koavalskaya Yu. M., Baskevich M. I. On species status of northern birch mice Sicista strandi Formosov (Rodentia, Dipodoidea). Zoologicheskii zhurnal, 1989, vol. 68, iss. 10, pp. 95–106 (in Russian).Sokolov V. E., Koavalskaya Yu. M., Baskevich M. I. On species status of northern birch mice Sicista strandi Formosov (Rodentia, Dipodoidea). Zoologicheskii zhurnal, 1989, vol. 68, iss. 10, pp. 95–106 (in Russian).Sumner A. T. A simple technique for demonstrating centromeric heterochromatin. Experimental Cell Research, 1972, vol. 75, iss. 1, pp. 304–306. https://doi.org/10.1016/0014-4827(72)90558-7Sumner A. T. A simple technique for demonstrating centromeric heterochromatin. Experimental Cell Research, 1972, vol. 75, iss. 1, pp. 304–306. https://doi.org/10.1016/0014-4827(72)90558-7Tamura K., Stecher G., Peterson D., Filipski A., Kumar S. MEGA6: Molecular Evolutionary Genetics Analysis version 6.0. Molecular Biology and Evolution, 2013, vol. 30, iss. 12, pp. 2725–2729. https://doi.org/10.1093/molbev/mst197Tamura K., Stecher G., Peterson D., Filipski A., Kumar S. MEGA6: Molecular Evolutionary Genetics Analysis version 6.0. Molecular Biology and Evolution, 2013, vol. 30, iss. 12, pp. 2725–2729. https://doi.org/10.1093/molbev/mst197Tsvetkova A. A., Oparin M. L. Biotopical expansion of small mammals in the steppe zone of the Zavolzhye. Arid Ecosystems, 2016, vol. 6, iss. 1, pp. 32–39. https://doi.org/10.1134/S2079096115040149Tsvetkova A. A., Oparin M. L. Biotopical expansion of small mammals in the steppe zone of the Zavolzhye. Arid Ecosystems, 2016, vol. 6, iss. 1, pp. 32–39. https://doi.org/10.1134/S2079096115040149Vinogradov B. S. Jerboas. Fauna of the USSR. Mammals. Vol. 3. Moscow, Leningrad, Academy of Sciences of the USSR Publ., 1937. 198 p. (in Russian).Vinogradov B. S. Jerboas. Fauna of the USSR. Mammals. Vol. 3. Moscow, Leningrad, Academy of Sciences of the USSR Publ., 1937. 198 p. (in Russian).Volga region. In: Bol'shaya Rossiiskaya ehntsiklopediya: v 30 t. [Big Russian Encyclopedia: in 30 vols]. Moscow, Bol'shaya Rossiiskaya ehntsiklopediya, 2014, vol. 26, pp. 24–26 (in Russian).Volga region. In: Bol'shaya Rossiiskaya ehntsiklopediya: v 30 t. [Big Russian Encyclopedia: in 30 vols]. Moscow, Bol'shaya Rossiiskaya ehntsiklopediya, 2014, vol. 26, pp. 24–26 (in Russian).Yasyuk V. P. Relikty v faune Samarskoi oblasti [Relics in the Fauna of the Samara Region]. Samara, Volga State Academy of Social Sciences and Humanities Publ., 2009. 104 p. (in Russian).Yasyuk V. P. Relikty v faune Samarskoi oblasti [Relics in the Fauna of the Samara Region]. Samara, Volga State Academy of Social Sciences and Humanities Publ., 2009. 104 p. (in Russian).

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